Znanstveno usavršavanje:

• 2009, 2010 - The Volcani Center, Institute for Plant Protection (Department of Entomology) Izrael, Laboratorij za molekularnu biologiju kukaca i genomiku, 7 mjeseci
• 2010 Advances in insect symbiosis research, International training school, COST FA0701, Hebrew University of Jerusalem, Rehovot, Israel
• 2010 Arthropod symbiont genomics and metagenomics, workshop, COST FA0701, University of Madeira, Funchal, Madeira (Portugal)
• 2008 In situ hybridization to plant chromosomes - a practical introduction to its uses in biosystematics, Prof. Leitch, Dr. Leitch, International graduate course, Croatian biological society; Faculty of Science, University of Zagreb, Croatia